Si tratta della sottospecie nemoralis legata ad ambienti più asciutti rispetto alla sottospecie nominale. Presente in poche stazioni tra Trentino, Veneto e Friuli.
I visited the serpentine quarry in Burgenland, Austria, where the sole, small population of Paragymnopteris marantae had been detected in 1962, but eradicated just 30 years later by widening the quarry.
My first sightings of the abundant Serpentine Spleenwort are likely close to the blasted away serpentine rocks which had been home to the most rare "Fur-fern" at northern border of it's range.
My trip went around the quarry and farther in the woods above. I could not find any site with Paragymnopteris marantae, so this is obviously gone lost there.
A. cuneifolium is obviously best capable to colonize fresh cracks in serpentine rocks as well as talus, so it is abundant around the quarry border. In opposite to that P. marantae seems to need boulders with steep flanks, which are relatively long exposed to withering, and this fern does likely propagate rather slowly, so there was no chance to survive at other, newly exposed rocks near by.